Stats for order Testudines

Stats for total mtDNA (group size: 34 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.8 - 41.5% 58.5 - 64.2% 3937 - 5122 1928 - 2467 3840 - 5518 5426 - 6734 14.7 - 127.0
Median 38.7% 61.3% 4418 2119 4487 5862 29.3
Average ± stdev 38.7 ± 1.1% 61.3 ± 1.1% 4469 ± 264 2113 ± 113 4521 ± 364 5898 ± 295 35.2 ± 21.7
Coefficient of variation 0.03 0.02 0.06 0.05 0.08 0.05 0.62
Pearson correlation
coefficient with log10MLS
0.01 -0.02 0.19 -0.05 0.06 0.17 -
Pearson p-value
(two tailed)
9.35E-01 9.13E-01 2.80E-01 7.90E-01 7.40E-01 3.50E-01 -
Stats for total protein-coding genes (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.7 - 42.8% 57.2 - 64.3% 2825 - 3575 1189 - 1455 2587 - 3283 3540 - 4021 14.7 - 127.0
Median 39.6% 60.4% 3195 1299 3022 3856 27.3
Average ± stdev 39.5 ± 1.3% 60.5 ± 1.3% 3189 ± 139 1294 ± 67 3015 ± 134 3845 ± 106 35.1 ± 22.0
Coefficient of variation 0.03 0.02 0.04 0.05 0.04 0.03 0.63
Pearson correlation
coefficient with log10MLS
0.08 -0.08 0.17 -0.19 -0.18 0.11 -
Pearson p-value
(two tailed)
6.55E-01 6.59E-01 3.51E-01 2.90E-01 3.14E-01 5.44E-01 -
Stats for D-loop (group size: 23 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 25.3 - 35.8% 64.2 - 74.7% 183 - 366 99 - 229 267 - 631 296 - 510 14.7 - 70.3
Median 31.0% 68.3% 209 121 401 368 29.3
Average ± stdev 31.0 ± 3.2% 68.7 ± 3.0% 228 ± 44 126 ± 25 408 ± 84 383 ± 53 31.6 ± 13.0
Coefficient of variation 0.10 0.04 0.19 0.20 0.21 0.14 0.41
Pearson correlation
coefficient with log10MLS
0.07 -0.15 0.21 0.17 0.03 0.19 -
Pearson p-value
(two tailed)
7.42E-01 4.96E-01 3.47E-01 4.44E-01 8.87E-01 3.83E-01 -
Stats for total tRNA-coding genes (group size: 34 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 37.0 - 39.5% 60.5 - 63.0% 338 - 377 228 - 247 387 - 426 519 - 568 14.7 - 127.0
Median 38.2% 61.8% 354 238 412 547 29.3
Average ± stdev 38.3 ± 0.7% 61.7 ± 0.7% 354 ± 9 237 ± 4 409 ± 9 545 ± 10 35.2 ± 21.7
Coefficient of variation 0.02 0.01 0.03 0.02 0.02 0.02 0.62
Pearson correlation
coefficient with log10MLS
0.33 -0.32 0.18 0.17 -0.23 -0.21 -
Pearson p-value
(two tailed)
5.94E-02 6.51E-02 2.96E-01 3.28E-01 1.89E-01 2.42E-01 -
Stats for total rRNA-coding genes (group size: 34 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 36.6 - 41.0% 59.0 - 63.4% 526 - 645 408 - 449 511 - 618 925 - 1036 14.7 - 127.0
Median 39.7% 60.3% 593 426 563 993 29.3
Average ± stdev 39.6 ± 1.0% 60.4 ± 1.0% 593 ± 20 425 ± 9 564 ± 21 989 ± 22 35.2 ± 21.7
Coefficient of variation 0.02 0.02 0.04 0.02 0.04 0.02 0.62
Pearson correlation
coefficient with log10MLS
-0.08 0.08 -0.02 -0.04 0.05 0.11 -
Pearson p-value
(two tailed)
6.68E-01 6.62E-01 9.32E-01 8.08E-01 7.69E-01 5.38E-01 -
Stats for ATP6 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.5 - 43.0% 57.0 - 64.5% 181 - 237 51 - 74 150 - 207 221 - 252 14.7 - 127.0
Median 37.4% 62.6% 193 63 189 239 27.3
Average ± stdev 38.1 ± 1.8% 61.9 ± 1.8% 197 ± 13 62 ± 6 187 ± 12 235 ± 7 35.1 ± 22.0
Coefficient of variation 0.05 0.03 0.07 0.10 0.07 0.03 0.63
Pearson correlation
coefficient with log10MLS
0.24 -0.24 0.26 -0.08 -0.27 0.09 -
Pearson p-value
(two tailed)
1.76E-01 1.76E-01 1.43E-01 6.49E-01 1.35E-01 6.26E-01 -
Stats for ATP8 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 29.1 - 39.5% 60.5 - 70.9% 38 - 58 6 - 15 29 - 60 62 - 79 14.7 - 127.0
Median 35.1% 64.9% 49 11 42 69 27.3
Average ± stdev 34.7 ± 2.3% 65.3 ± 2.3% 49 ± 4 10 ± 2 42 ± 6 69 ± 4 35.1 ± 22.0
Coefficient of variation 0.07 0.04 0.09 0.23 0.16 0.06 0.63
Pearson correlation
coefficient with log10MLS
0.10 -0.10 0.32 -0.24 -0.12 0.25 -
Pearson p-value
(two tailed)
5.69E-01 5.69E-01 7.01E-02 1.71E-01 4.98E-01 1.56E-01 -
Stats for COX1 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 38.0 - 43.5% 56.5 - 62.0% 348 - 427 218 - 276 414 - 493 426 - 484 14.7 - 127.0
Median 41.3% 58.7% 383 252 459 454 27.3
Average ± stdev 41.2 ± 1.2% 58.8 ± 1.1% 385 ± 18 251 ± 13 454 ± 18 453 ± 14 35.1 ± 22.0
Coefficient of variation 0.03 0.02 0.05 0.05 0.04 0.03 0.63
Pearson correlation
coefficient with log10MLS
-0.00 0.01 0.15 -0.22 -0.17 0.23 -
Pearson p-value
(two tailed)
9.82E-01 9.59E-01 4.09E-01 2.29E-01 3.45E-01 2.01E-01 -
Stats for COX2 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 34.5 - 41.6% 58.4 - 65.5% 149 - 202 77 - 103 153 - 208 228 - 255 14.7 - 127.0
Median 38.1% 61.9% 173 87 182 242 27.3
Average ± stdev 37.9 ± 1.7% 62.1 ± 1.7% 172 ± 12 88 ± 5 182 ± 11 243 ± 6 35.1 ± 22.0
Coefficient of variation 0.05 0.03 0.07 0.06 0.06 0.02 0.63
Pearson correlation
coefficient with log10MLS
0.12 -0.12 0.11 -0.01 -0.17 0.10 -
Pearson p-value
(two tailed)
4.94E-01 5.07E-01 5.24E-01 9.72E-01 3.32E-01 5.67E-01 -
Stats for COX3 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 39.2 - 46.7% 53.3 - 60.8% 197 - 240 99 - 130 187 - 229 224 - 258 14.7 - 127.0
Median 41.9% 58.1% 211 116 214 241 27.3
Average ± stdev 42.0 ± 1.4% 58.0 ± 1.4% 213 ± 9 115 ± 6 212 ± 9 241 ± 7 35.1 ± 22.0
Coefficient of variation 0.03 0.02 0.04 0.06 0.04 0.03 0.63
Pearson correlation
coefficient with log10MLS
0.22 -0.22 0.15 0.06 -0.21 -0.15 -
Pearson p-value
(two tailed)
2.20E-01 2.20E-01 4.15E-01 7.53E-01 2.39E-01 4.04E-01 -
Stats for CYTB (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 37.7 - 45.6% 54.4 - 62.3% 301 - 375 119 - 145 266 - 332 301 - 381 14.7 - 127.0
Median 42.0% 58.0% 347 131 305 359 27.3
Average ± stdev 41.9 ± 1.6% 58.1 ± 1.6% 346 ± 16 131 ± 5 304 ± 13 357 ± 16 35.1 ± 22.0
Coefficient of variation 0.04 0.03 0.05 0.04 0.05 0.05 0.63
Pearson correlation
coefficient with log10MLS
0.08 -0.08 0.13 -0.20 -0.22 0.07 -
Pearson p-value
(two tailed)
6.52E-01 6.52E-01 4.70E-01 2.76E-01 2.13E-01 6.87E-01 -
Stats for ND1 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 36.4 - 43.5% 56.5 - 63.6% 252 - 317 99 - 129 240 - 298 275 - 331 14.7 - 127.0
Median 39.5% 60.5% 274 107 274 315 27.3
Average ± stdev 39.6 ± 1.5% 60.4 ± 1.5% 275 ± 13 108 ± 6 274 ± 12 312 ± 11 35.1 ± 22.0
Coefficient of variation 0.04 0.03 0.05 0.06 0.05 0.04 0.63
Pearson correlation
coefficient with log10MLS
0.15 -0.15 0.22 -0.10 -0.10 -0.06 -
Pearson p-value
(two tailed)
4.17E-01 4.17E-01 2.22E-01 5.70E-01 5.75E-01 7.60E-01 -
Stats for ND2 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 33.8 - 40.6% 59.4 - 66.2% 257 - 324 77 - 123 235 - 289 355 - 409 14.7 - 127.0
Median 38.6% 61.4% 307 94 257 382 27.3
Average ± stdev 38.3 ± 1.5% 61.7 ± 1.5% 303 ± 15 94 ± 9 259 ± 12 382 ± 12 35.1 ± 22.0
Coefficient of variation 0.04 0.02 0.05 0.10 0.05 0.03 0.63
Pearson correlation
coefficient with log10MLS
0.08 -0.08 0.31 -0.37 -0.19 0.09 -
Pearson p-value
(two tailed)
6.55E-01 6.55E-01 7.77E-02 3.22E-02 3.01E-01 6.04E-01 -
Stats for ND3 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 33.8 - 43.3% 56.7 - 66.2% 79 - 109 31 - 47 80 - 121 93 - 125 14.7 - 127.0
Median 38.3% 61.7% 92 39 106 113 27.3
Average ± stdev 38.1 ± 2.2% 61.9 ± 2.2% 94 ± 7 38 ± 4 102 ± 8 113 ± 6 35.1 ± 22.0
Coefficient of variation 0.06 0.04 0.08 0.11 0.08 0.05 0.63
Pearson correlation
coefficient with log10MLS
0.30 -0.30 0.19 0.13 -0.25 -0.13 -
Pearson p-value
(two tailed)
8.60E-02 8.60E-02 2.82E-01 4.83E-01 1.57E-01 4.79E-01 -
Stats for ND4 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.0 - 43.3% 56.7 - 65.0% 348 - 463 119 - 169 292 - 395 448 - 513 14.7 - 127.0
Median 38.9% 61.1% 383 139 367 484 27.3
Average ± stdev 38.5 ± 1.8% 61.5 ± 1.8% 388 ± 23 142 ± 14 366 ± 19 481 ± 17 35.1 ± 22.0
Coefficient of variation 0.05 0.03 0.06 0.10 0.05 0.04 0.63
Pearson correlation
coefficient with log10MLS
-0.20 0.20 0.02 -0.32 0.01 0.33 -
Pearson p-value
(two tailed)
2.56E-01 2.56E-01 9.33E-01 6.61E-02 9.59E-01 5.68E-02 -
Stats for ND4L (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.0 - 44.1% 55.9 - 65.0% 78 - 103 23 - 39 72 - 101 82 - 104 14.7 - 127.0
Median 40.1% 59.9% 87 32 88 89 27.3
Average ± stdev 40.1 ± 2.0% 59.9 ± 2.0% 87 ± 5 31 ± 3 88 ± 5 89 ± 5 35.1 ± 22.0
Coefficient of variation 0.05 0.03 0.06 0.13 0.06 0.06 0.63
Pearson correlation
coefficient with log10MLS
-0.00 0.00 -0.02 -0.03 0.04 -0.10 -
Pearson p-value
(two tailed)
9.88E-01 9.88E-01 9.11E-01 8.88E-01 8.17E-01 5.93E-01 -
Stats for ND5 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 34.5 - 41.9% 58.1 - 65.5% 447 - 565 163 - 215 406 - 518 595 - 666 14.7 - 127.0
Median 39.0% 61.0% 525 181 467 626 27.3
Average ± stdev 38.9 ± 1.3% 61.1 ± 1.3% 518 ± 24 181 ± 11 470 ± 27 628 ± 17 35.1 ± 22.0
Coefficient of variation 0.03 0.02 0.05 0.07 0.06 0.03 0.63
Pearson correlation
coefficient with log10MLS
0.04 -0.04 0.01 0.07 -0.09 0.10 -
Pearson p-value
(two tailed)
8.30E-01 8.30E-01 9.56E-01 6.93E-01 6.24E-01 5.76E-01 -
Stats for ND6 (group size: 33 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 29.9 - 44.6% 55.4 - 70.1% 130 - 184 27 - 48 52 - 108 204 - 260 14.7 - 127.0
Median 38.3% 61.7% 162 42 76 249 27.3
Average ± stdev 38.3 ± 2.4% 61.7 ± 2.4% 160 ± 9 40 ± 5 76 ± 8 247 ± 10 35.1 ± 22.0
Coefficient of variation 0.06 0.04 0.06 0.13 0.11 0.04 0.63
Pearson correlation
coefficient with log10MLS
0.16 -0.16 0.21 -0.03 -0.17 -0.03 -
Pearson p-value
(two tailed)
3.85E-01 3.85E-01 2.35E-01 8.59E-01 3.30E-01 8.48E-01 -
Stats for 12S rRNA gene (group size: 34 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 38.6 - 43.5% 56.5 - 61.4% 211 - 254 156 - 182 196 - 229 338 - 373 14.7 - 127.0
Median 41.4% 58.7% 237 164 211 357 29.3
Average ± stdev 41.3 ± 1.0% 58.7 ± 1.0% 236 ± 8 164 ± 4 211 ± 7 357 ± 7 35.2 ± 21.7
Coefficient of variation 0.02 0.02 0.04 0.03 0.04 0.02 0.62
Pearson correlation
coefficient with log10MLS
0.14 -0.14 0.18 0.02 -0.13 0.00 -
Pearson p-value
(two tailed)
4.14E-01 4.26E-01 3.06E-01 8.91E-01 4.65E-01 9.98E-01 -
Stats for 16S rRNA gene (group size: 34 species)
GC (%) AT (%) G (bases) C (bases) A (bases) T (bases) Maximum lifespan (yrs)
Range (min-max) 35.3 - 40.3% 59.7 - 64.7% 315 - 391 249 - 275 314 - 389 587 - 663 14.7 - 127.0
Median 38.6% 61.4% 360 262 350 635 29.3
Average ± stdev 38.6 ± 1.1% 61.4 ± 1.1% 356 ± 15 261 ± 6 353 ± 16 631 ± 16 35.2 ± 21.7
Coefficient of variation 0.03 0.02 0.04 0.02 0.05 0.03 0.62
Pearson correlation
coefficient with log10MLS
-0.18 0.18 -0.12 -0.08 0.13 0.15 -
Pearson p-value
(two tailed)
3.08E-01 3.08E-01 4.96E-01 6.45E-01 4.71E-01 3.95E-01 -

Contact: Vadim E. Fraifeld, MD, PhD

Head: Lab for the Biology of Aging, The Shraga Segal Department of Microbiology, Immunology and Genetics, Faculty of Health Sciences, Ben-Gurion University of the Negev
P.O. Box 653, Beer-Sheva 8410501, Israel

Email:
How to cite us:

If you would like to cite this database please use:
Toren D, Barzilay T, Tacutu R, Lehmann G, Muradian KK, Fraifeld VE. MitoAge: a database for comparative analysis of mitochondrial DNA, with a special focus on animal longevity. Nucleic Acids Res. 2016; 44(D1):D1262-5.